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Llantwit Major — Llanilltud Fawr — is one of the oldest continuously inhabited places in Wales, an ancient monastic town set back a mile from the Heritage Coast. Its medieval church, old town lanes, and surrounding farmland retain an unusually undisturbed character. The cliff path between Llantwit Beach and Ogmore-by-Sea to the west is one of the finest sections of the Glamorgan Heritage Coast, lined with bramble, gorse, thrift, and cliff-top grassland that supports bees well into September. The old farm lanes between Llantwit Major and Boverton have the hedgerow hawthorn and elder typical of the inner Vale.

Blackthorn and cherry plum open the year in the old orchards and thickset hedgerows around Cowbridge and Llantwit Major — some of the most intact ancient-enclosure hedge networks remaining in Wales. Hawthorn follows through the early Vale fields. White clover still dominates the traditionally managed meadows between Rhoose and St Athan, and oilseed rape is grown sporadically on the lighter soils. The June highlight is lime — Barry, Penarth, and Cowbridge all have fine street limes and park limes — followed by a long bramble flow along the Heritage Coast cliff paths. Sycamore is useful on the sheltered coastal slope; sea buckthorn, thrift, and bird's-foot trefoil supplement on the cliff grassland. Ivy on limestone walls and old farmsteads closes a long, gentle season.